Macronutrients (N, P, K, Ca, Mg, and S) in litter of three primarily spruce (Picea purpurea Masters) (SF), fir (Abies faxoniana Rehder & E. H. Wilson) (FF), and birch (Betula platyphylla Sukaczev) (BF) subalpine forests in western China were measured to understand the monthly variations in litter nutrient concentrations and annual and monthly nutrient returns via litterfall. Nutrient concentration in litter showed the rank order of Ca > N > Mg > K > S > P. Monthly variations in nutrient concentrations were greater in leaf litter (LL) than other litter components. The highest and lowest concentrations of N, P, K, and S in LL were found in the growing season and the nongrowing season, respectively, but Ca and Mg were the opposite. Nutrient returns via litterfall showed a marked monthly pattern with a major peak in October and one or two small peaks in February and/or May, varying with the element and stand type, but no marked monthly variations in nutrient returns via woody litter, reproductive litter, except in May for the BF, and moss litter. Not only litter production but also nutrient concentration controlled the annual nutrient return and the monthly nutrient return pattern. The monthly patterns of the nutrient concentration and return were of ecological importance for nutrient cycling and plant growth in the subalpine forest ecosystems.
Litter production, components and dynamics were investigated and forest floor litter was quantified throughout awhole year in three subalpine forests, dominated by tree species of spruce (SF), fir (FF) and birch (BF), in WesternSichuan, China, in order to understand the key factors that influenced litter production and dynamics. Litterfall in thethree forests consisted mainly of leaves, woody litter, reproductive organs and moss. Contribution of leaf litter to thetotal litterfall was significantly (P < 0.05) greater than that of woody litter, reproductive organs or moss. Regardlessof the stands, litterfall exhibited a marked monthly variation with the maximum litterfall peaks occurring in October,with smaller peaks occurring in February for SF and FF, and May for BF. The analysis indicated that tree species,stand density, leaf area index (LAI), stand basal area and stand age were the key factors determining litter production.Meanwhile tree species and phenology controlled the litter dynamics, with wind and snow modifying the litter componentsand dynamics.