Three new species of fossil Palaeontinidae are described from Daohugou Village, Inner Mongolia, China: Daohugoucossus shii sp. nov., D. parallelivenius sp. nov., D. lii sp. nov. The diagnosis of Dao- hugoucossus Wang, Zhang and Fang, 2006 is revised. These materials are the first complete fossil palaeontinids in the Middle Jurassic of the world. Based on observation of these new specimens, Sc of this new genus has the following combined characters: joined with R at base, diverged near M separa- tion, coalesced with R before the nodal line, extended beyond the nodal line and fused with R1 before distal end. Due to the Sc characteristics mentioned above, the genus Liaocossus Ren, Yin and Dou, 1998 can be well distinguished from “Ilerdocossus-complex”.
One new genus and three new species of the weevil family Eccoptarthridae (Curculionoidea), Leptocar polychaetus gen. et sp. nov., Abrocar macilentus sp. nov., Cretonanophyes punctatus sp. nov., are described and illustrated. They pertain to the Upper Jurassic or Lower Cretaceous Yixian Formation of western Liaoning Province, China. The host plants to the living eccoptarthrids and the floras in Yixian Formation indicate that those archaic eccoptarthrids possibly lived on conifers with a phylogenetic closeness to Cupressaceae. Besides, the early diversification of eccoptarthrids is discussed: origi- nated in Central or East Asia in the Late Jurassic, spread into Western Europe and South America during the Early Cretaceous, underwent a dramatic decline after the Early Cretaceous which likely was caused by competitive pressure and the displacement of their host plants.
